- Directors: David Frankel
- Producers: Wendy Finerman, Karen Rosenfelt
- Writers: Aline Brosh McKenna, Novel, Lauren Weisberger
- Genres: Comedy, Drama, Romance
- Actors: Anne Hathaway, Meryl Streep, Emily Blunt, Stanley Tucci, Simon Baker, Adrian Grenier
Andrea “Andy” Sachs (Anne Hathaway), an aspiring journalist fresh out of Northwestern University, lands the magazine job “a million girls would kill for”: junior personal assistant to icy editor-in-chief Miranda Priestly (Meryl Streep), who dominates the fashion world from her perch atop Runway magazine. She puts up with the eccentric and humiliating requests of her boss because, she is told, if she lasts a year in the position she will get her pick of other jobs, perhaps even the journalistic position she truly craves.
At first, she fits in poorly among the gossipy fashionistas who make up the magazine staff. Her lack of style or fashion knowledge and fumbling with her job make her an object of scorn around the bustling office. Senior assistant Emily Charlton (Emily Blunt), her co-worker, is condescending to her. Gradually, though, with the help of art director Nigel (Stanley Tucci), Andy adjusts to the position and its many perks, including free designer clothing and other choice accessories. She begins to dress more stylishly and do her job competently, fulfilling a seemingly impossible request of Miranda’s to get two copies of an unpublished Harry Potter manuscript to her daughters.
She also comes to prize chance encounters with attractive young writer Christian Thompson (Simon Baker), who helped her obtain the Potter manuscript and suggests he could help her with her career. Her relationships with her boyfriend Nate (Adrian Grenier), a chef working his way up the career ladder, and other college friends begin to suffer due to the increasing time she spends at Miranda’s beck and call.
Later, back in New York, she meets Nate for breakfast. He has accepted an offer to work as a sous-chef in a popular Boston restaurant. Andrea is disappointed, but her hope is rejuvenated when he says they could work something out. At the film’s conclusion, she is interviewing for a newspaper job. The interviewer reveals that he received a fax from Miranda saying she was by far her biggest disappointment, but that if he did not hire her, he was an idiot. Andrea offers Emily the clothes she wore to Paris, insisting she doesn’t need them anymore. Emily accepts and tells Andrea’s replacement she has some large shoes to fill. In the last scene, Andrea, dressed casually but with a bit more style, sees Miranda getting into her car across the street. They exchange looks and Miranda gives a soft smile once inside the car. She then snaps back to her usual self and impatiently says “Go!” to the driver.